This is part of the #1796 series. Read that issue for context
Test source
Ref source (mismatch)
Live test
Live ref (mismatch)
As seen in the test above, in the general case, if a inline-level box contains only characters from fallback fonts, all browsers agree to size the height of a line-height:auto inline-level box using the metrics of the fonts actually used plus the metrics of the primary font, even if it is not used. I.E. they take the strut into account.
However, see #1798 for a possible exception.